ANTS
 
Several species of ants commonly inhabit home lawns and ornamental plantings of trees and shrubs. Winged ants are frequently confused with swarming termites. Ants can be easily distinguished from termites by the following characteristics: (1) ants have elbowed antennae while those of termites are straight and beadlike; (2) front wings of ants are larger in size than the rear wings while those of termites are all equal in size; (3) the waist of ants are thin and appear to be constricted while those of termites are broad and not constricted; (4) wings of ants are usually transparent or brownish while those of termites are milky-white.
 
DESCRIPTION
 
Little Black Ant: Workers of this species are 1/10-1/8 inch in length with soft jet-black bodies. These ants are found primarily in soil and rotting wood. They feed on a wide variety of food sources.

The Pavement Ant: These ants are slow, sluggish, short-legged, and usually nest under pavements and foundations. They are 1/12-1/4 inch in length and brownish-black. They feed on a wide variety of food sources (seeds, grease, animal food).

Cornfield Ant: These ants are usually found nesting in open places in the soil or in rotten wood. They are 1/8-1/4 inch in length, robust, soft-bodied, and light to dark brown. The body, when crushed, has an acid (formic) odor. These ants are found feeding on seeds, and often are associated with aphids and mealybugs which are excreting honeydew.

Larger Yellow Ant: These ants are 1/10-1/8 inch in length and yellow. When crushed they give off a characteristic lemon-verbena odor. They are common soil inhabiting species which tend mealybugs and aphids on the roots of plants.

LIFE HISTORY

Ants form colonies or nests where queens remain. Most ant species have only one queen per nest; she lays the eggs needed to maintain or increase the colony. Queens may live from 1-15 years and produce many thousands of eggs.

Larvae, or maggot-like grubs, hatch from the eggs. They are transparent white, soft-bodied, and legless. These larvae, along with the queen, are fed by the worker ants (foragers). Larvae pass through several molts before pupation and adulthood are reached. The majority of these develop into workers; a few develop into winged males and females which are the kings and queens of the ant colony. At certain times during the year (usually early spring and late summer), varying with the species, these winged ants leave the nest and swarm. Females and males mate and the males die soon after. The mated female (queen) flies to an attractive nesting site, tears off her wings, and encloses herself in a small excavation in the soil, and lays eggs. She feeds and cares for the first generation of progeny until they are mature adults. After that they and succeeding generations care for her. She remains in the nest and continues to produce eggs the remainder of her life.

DAMAGE

Ants seldom cause serious damage to home lawns. They may cause minor damage to the turf by loosening the soil and constructing small mounds in the nesting areas. Ants also feed on honeydew (sticky sugary substance) excreted by aphids, mealybugs, and soft scales. Homeowners may become alarmed when ants swarm during spring and late summer because they associate swarming insects with termites. However, if they examine specimens for the above differentiating characters, confusion should be eliminated. Respect these small insects since they do bite and some can also sting. Many ants are considered beneficial because they are scavengers and predators on certain harmful insects.

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Armyworm

Native to North America, the armyworm can be found east of the Rocky Mountains, reaching northward into southern Canada. Armyworms get their name from their behavior of moving across fields in an army-like fashion. As larvae consume available food sources, they migrate as an army to new host plants. Though they feed primarily on grasses (oats, wheat, fall rye, corn, barley, and forage grasses), they can be a pest of some vegetables (bean, cabbage, carrot, onion, pea, pepper, radish and sweet potato). There are usually two to three generations each year in Illinois.

Armyworm Larva
Armyworm larva

Description

Armyworm larvae vary in color from dark greenish-brown to black. On each side, there are long, pale white, orange, and dark brown stripes along the length of the abdomen. Mature larvae are approximately 1 ˝ inches long. The head capsule is yellowish brown with a brown network of veins, giving it a mottled appearance. Armyworm larvae may also be distinguished by a dark band on the outer side of each proleg. Armyworm pupa stay in a brown earthen shell just below the soil surface. The armyworm moth, approximately an inch long with a 1 ˝ inch wingspan, is tan to light brown, with a tiny white spot centered on each forewing. Eggs, which resemble small white globules, are laid in rows or groups on leaves of host plants. After oviposition, the moth rolls the leaf blade of the plant around each egg mass.

Life Cycle

Few armyworms overwinter in Illinois. Most armyworm moths migrate into the Illinois from the southern states in April and May. During the day, moths remain hidden in grassy vegetation. Armyworm moths are active during the evening, feeding on nectar, mating, and searching for oviposition sites. Eggs are deposited in rows or clusters on thelower leaves of grasses or at the base of plants. Eggs hatch in1 to 2 weeks. Newly hatched larvae are pale green and move in a looping motion. Larvae are also active at night, feeding on host plants. During the day, they can be found under plant debris or in the top few inches ofthe soil. After completing six instars, larvae pupate just below the soil surface. Adults emerge in 1 to 2 weeks. A second generation occurs in late June or early July and a third in late August or early September.

Injury

Since moths prefer dense vegetation for oviposition sites, armyworm infestations generally develop in areas such as grass pastures, roadsides, and along fence rows. Oviposition sites often include weeds and grassy in weedy or reduced tillage fields. Consequently, these fields armyworms are often a problem in these fields. If a herbicide is used to control the weeds, larvae move from the dead grasses to the corn. As larvae run out of available food sources, they move to other host plants such as small grains and corn. This usually occurs during May and early June.

Armyworms consume leaf tissue of corn plants. Feeding is usually confined to leaf margins, but in some instances, larvae may strip the plants entirely of leaf tissue. Corn generally recovers from damage caused by moderate infestations if the growing pointhas not been injured. Ordinarily armyworms that attack young corn migrate from small grains or grass fields.

Armyworm larvae chew the leaves of small grains and grasses. They may strip the leaf margins and move up the plant to feed on the panicles or flowers. Larvae will feed on the flag leaves, kernels, and clip the stems just below the heads.

Injury caused by armyworms in forages is sometimes confused with that caused by other insects. Armyworms do not attack pure stands of alfalfa and other legumes, but cutworms do feed on these crops. If the field is a grass-legume mixture, cutworms and armyworms may both be present. If it is a pure grass stand, the insects are probably armyworms. The damage is the same, however, as both eat the foliage.

Oftentimes groundhogs, rabbits, mice, and other small animals will cut stems of plants into small sections and pile them up. All of the plants in a small area may be cut up into tiny sections. Many people mistake this for armyworm damage.

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Billbugs

David J. Shetlar

Billbugs are weevils that have the snout, head and thorax about as long as the wing covers. Though there are about a half dozen species that may be found in turfgrasses, only a few cause damage in any given region. The most common pest in Ohio is the bluegrass billbug, Sphenophorus parvulus Gyllenhal, though the lesser billbug, S. minimus Hart, is somewhat common. The adults are only 1/4- to 3/8-inch long and dark grey to black. They may be covered with a tan or brown coating of soil. The larvae are white with a brown head and look like legless white grubs.
Figure 1. Bluegrass Larva

The bluegrass billbug is most common in the northern states from New England to Washington state. It is found less commonly in southern states. Both billbugs seem to prefer Kentucky bluegrass-growing regions.
Figure 2. Bluegrass Billbug Adult

Kentucky bluegrass seems the preferred host but occasionally this pest has infested perennial ryegrass, red fescue and tall fescue. It also attacks small grain crops such as corn, rye and wheat.

Types Of Damage

Billbug damage usually appears in late-June through August, when summer drought stress is common. Light infestations in lawns often produce small dead spots that look like the turf disease, dollar spot. Sometimes the damage looks like irregular mottling or browning in the turf. Heavy infestations can result in complete destruction of the turf, usually by August. The major problem with billbug damage is that it looks like a variety of other problems. Most turf managers confuse billbug damage with drought, disease, chinch bugs, greenbugs or white grubs. Billbug damaged turf turns a whitish-straw color rather than the yellow caused by greenbugs. Soil under damaged turf is solid, not spongy as in white grub attacks.

To confirm billbug attacks, grasp the affected turf and pull upward. If the stalks break easily at ground level and the stems are hollowed out or are full of packed sawdust-like material, billbugs are the culprit.

Life Cycle And Habits

In most of the Kentucky bluegrass growing regions this pest overwinters in the adult stage. Adults have been found in thatch, cracks and crevices in the soil, worm holes and in leaf litter near turf. The hibernating adults become more active in late-April to mid-May when the soil surface temperatures rise above 65 degrees F. The adults wander in search of suitable grasses and crops on which to feed. After feeding for a short period, the female begins to insert one to three eggs in a feeding hole made in grass stems. The females may continue laying eggs into August but most eggs are laid by early-July. Laboratory kept females have been known to lay over 200 eggs, usually two to five per day. The eggs hatch in six days depending on the temperature and the young larvae begin to tunnel up and down the stem. If a stem is hollowed out while the larva is small, an exit hole may be formed and the larva will drop out and bore into another stem. Eventually the larva becomes too large to fit inside the grass stems. They then drop to the ground to begin feeding externally on the grass crowns and roots. This is the point at which significant damage to the turf is noticed, especially if little rainfall or irrigation has occurred at this time. After 35 to 55 days, the larva is full grown and pupates in a cell of soil under the thatch. The pupa gradually darkens and the reddish-brown, tineral adult emerges in 8 to 10 days. The new adults appear to be common in late-August through September. These adults do some minor feeding and seek out suitable sites for overwintering. Some adults have been observed trying to fly but no great distances were covered. There is some evidence that adults which emerge early in August may begin laying eggs for a partial second generation. These larvae often do not develop rapidly enough to mature before freezing temperatures arrive.

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Chinch Bugs

David J. Shetlar

There are several chinch bugs that attack turfgrasses in North America. The hairy chinch bug, Blissus leucopterus hirtus Montandon, is the most commonly encountered pest of northern turfgrasses though the common chinch bug, Blissus leucopterus leucopterus Say, is occasionally found. These closely related pests are very difficult to separate in the field and most people identify them by locality and type of food plants.

The common chinch bug is normally found from South Dakota across to Virginia and south to a line running from mid-Texas across to mid-Georgia. The hairy chinch bug cohabits some of the northern range of the common chinch bug but also extends throughout the northeastern states and into southern Canada.

The hairy chinch bug prefers turfgrass species such as fine fescues, perennial ryegrasses, Kentucky bluegrass, bentgrass and zoysiagrass. The common chinch bug prefers grain crops such as sorghum, corn and wheat but will attack turfgrasses such as Bermudagrass, fescues, Kentucky bluegrass, perennial ryegrass, zoysiagrass and crabgrass.
Shortwinged Adult Chinch Bugs Bigeyed Chinch Bugs
Shortwinged Adult Bigeyed Bug Adult

Types of Damage

Chinch bug damage is usually first detected when irregular patches of turf begin to turn yellow then straw colored. The straw colored areas may be completely dead. These patches continue to become larger in spite of watering.

Apparently, feeding by chinch bugs blocks the water and food conducting vessels of grass stems. By blocking the water, the leaves wither as in drought and the manufactured food doesn't get to the roots. The result is plant death. Damage generally occurs during hot, dry weather from June into September.

Description of Stages

These pests are true bugs and have a gradual life cycle with egg, nymphal and adult stages. All the species of Blissus are very similar in form and an expert is needed to separate species and subspecies.

Eggs

The eggs are elongate, bean-shaped, approximately 0.84 mm long by 0.25 mm wide, and are roundly-pointed at one end and blunt at the other. The blunt end has several small tubercles visible through a dissecting microscope. The eggs are first white and change to bright orange just before hatching.

Nymphs

There are five nymphal instars which change considerably in color and markings. The first instar has a bright orange abdomen with a cream colored stripe across it, a brown head and thorax and is about 0.9 mm long. The second through fourth instars continue to have this same general color pattern except that the orange color on the abdomen gradually changes to a purple-gray with two black spots. The fourth instar increases to more than 2 mm long. The fifth instar is very different because the wing pads are easily visible and the general color is now black. The abdomen is blue-black with some darker black spots and the total body length is about 3 mm.

Adults

The adults are approximately 3.5 mm long and 0.75 mm wide. The males are usually slightly smaller than the females. The head, pronotum and abdomen are gray-black in color and covered with fine hairs. The wings are white with a black spot (the corium) located in the middle of the front-wing edge. The legs often have a dark burnt orange tint. Individuals in a population, or in some cases, most of a local population may have short, called brachypterous, wings which reach only half-way down the abdomen.

Chinch Bug Life Cycles

Life Cycles and Habits

The hairy chinch bug adults overwinter in the thatch and bases of grass stems in the turf. However, the common chinch bug prefers to move to tall bunch-grasses in open fields to find overwintering sites. These individuals then migrate in search of grain crops in the spring but may establish in turf instead. The adults of both species become active when the daytime temperatures reach 70 degrees F. The females feed for a short period of time and mate when males are encountered. Eventually the females begin to lay eggs by inserting them into the folds of grass blades or into the thatch. This usually occurs from mid-April into June, from New York to Illinois. A single female may lay up to 200 eggs over 60-80 days. The eggs take about 20-30 days to hatch at temperatures below 70 degrees F but can hatch in as little as a week when above 80 degrees F. The young nymphs begin to feed by inserting their mouthparts in grass stems, usually while under a leaf sheath. The nymphs grow slowly at the beginning of the season because of cool temperatures but speed their development by July. Usually the first generation matures by mid-July. At this time considerable numbers of adults and larger nymphs can be seen walking about on sidewalks or crawling up the sides of light colored buildings. If a good, hot, dry spring is available, turf injury by the first generation can be evident by June. Damage may be visible from late-June through August when the spring generation mature nymphs and adults are feeding and the second generation of nymphs are becoming active. During the hot summer months, the new females lay eggs rapidly and their young may mature by the end of August into September. The second generation adults may lay a few eggs for a partial third generation if the season has been long. However, most of these late nymphs do not mature before winter temperatures drop. When cool temperatures arrive, the mature chinch bugs seek out protected areas to spend the winter.

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Cutworms

The larvae or caterpillars of some moths are called cutworms (Agrotis, Amathes,Black cutworm Peridroma, Prodenia spp.) because of the manner in which they cut down young plants as they feed. The adults are night-flying moths which feed on nectar, if at all, and do no damage.

Damage:

There are a great many species of cutworms. While they all feed on plants by chewing, they vary as to damage done and host plants preferred. Generally they destroy more of the plant than they eat. Their numbers vary greatly from year to year and, when numerous, may destroy as much as 75% of a crop. Cutworms injure plants in four major ways:

- Solitary surface cutworms cut off young plants at or slightly above or below the soil line, sometimes dropping the severed plants into their burrows. Because most of the plant is not eaten, these cutworms do great damage, attacking and felling new plants nightly. The black, bronzed, clay-backed and dingy cutworms are in this group.

 

Black cutworm and damaged stalk
(Clemson University Extension)

  • Climbing species, usually the variegated and spotted cutworms, climb the stem of trees, shrubs, vines, and crops and eat the leaves, buds and fruit.
  • Subterranean species, particularly the pale western and glassy cutworms, remain in the soil and feed upon roots and underground parts.
  • Army cutworms occur in great numbers, consuming the tops of plants and then "marching" on to other fields.

Description:

The many species of cutworms can be quite distinct. Many are stout, smooth, soft-bodied, plump caterpillars. These vary from brown or tan to pink, green or gray and black. Some are all one color, others spotted or striped. Some larvae are dull, others appear glassy. The adults are generally very robust brown or black moths showing various splotches, blotches or stripes in shades of gray, brown, black or white. 

Life Cycle:

Most cutworms pass the winter as partially grown larvae. Thus they are already large, voracious feeders when transplants and seedlings are set out in the fields. A few species pass the winter as pupae or hibernating moths. Overwintering cutworms may live under trash or bark, in clumps of grass or in earthen cells in the soil. These cutworms become active and begin feeding as the weather warms in spring, remaining hidden under debris or in the soil and feeding at night. Many species continue to feed well through June, then pupate in the soil to emerge later as moths. Normally there is only one generation per year. The moths crawl from their brown pupal cases in the soil and climb up through the soil, following the tunnel made by the burrowing larva. If this tunnel is blocked, the fragile moth cannot escape the soil. Cutworm abundance and development is greatly affected by weather, especially rainfall. The moths mate and lay eggs in late summer, beginning the next generation. The moths often seek out grassy or weedy areas to lay their eggs, which are usually deposited on plant stems or in the soil. One female may lay hundreds of eggs. The hatching larvae feed until cold weather and then hide for the winter in a sheltered, dry place.

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Deer Tick: 
 
The most important pest to control in your backyard!

Facts To Know About Deer Ticks

The Deer Tick, Ixodes dammini, is the vector for Lyme disease in the northeastern United States, primarily because this tick has been most extensively studied. There is ongoing debate whether I. dammini is conspecific with I. scapularis, the member of the I. ricinus complex present in the southeastern United States. What is clear is that the northern tick population poses a much greater threat of transmitting Lyme disease. Possible explanations for the apparent increase in the population of I. dammini and the expansion of its geographic range are reviewed, as is the impact of changing patterns of land use in the United States over the past century, which have created environments ideally suited for ticks and deer (the preferred host for adult I. dammini ticks) and which bring humans into close proximity to both.

An important concept (albeit relatively simple) is that the total population of ticks in a region is a result of the balance among fecundity (the number of offspring produced per female), mortality, and migration. The maximal number of offspring per adult female tick is estimated to be approximately 3000.

In view of the tick's two-year life cycle, assuming equal sex distribution of progeny, a single female could produce up to 4.5 million offspring in just two years. This astounding fact suggests a tremendous excess tick mortality during development. However, little is known about how to increase this mortality so that the risk of Lyme disease can be meaningfully reduced. It also suggests that approaches focused on adult female ticks might be the more fruitful than approaches focused on ticks in immature stages.

Possible techniques of environmental management to reduce the risk of the disorder will require IPM. Options discussed include measures for personal protection as well as environmental measures, including "habitat modification" (e.g., burning underbrush), "host exclusion" (e.g., deer fencing), and both "off-host" methods (e.g., insecticides) and"on-host" options to increase tick mortality. Several possibilities of particular scientific interest are distributing insecticide-impregnated cotton balls in the environment where mice (important hosts forimmature I. dammini ticks) may use them in nesting material; developing vaccines against tick saliva to promote rejection of the tick during feeding, thereby preventing engorgement, particularly of adults feeding on deer, and thus interrupting the tick's life cycle before mating; and developing mixtures of pheromones with acaricides. Pheromones are chemical compounds produced by ticks as chemical signals to facilitate communication. Some pheromones are attractants (for example, sex pheromones); they could be tagged with tick insecticide (acaricide), thus increasing its power.

It is highly likely that the successful control of Lyme disease will require an integrated management approach that will include personal-protection measures, improved diagnostic and therapeutic approaches, and environmental interventions at many levels.

In this era of increased consciousness of the costs of public health as well as those of individual health care, interventions must be chosen with attention to what is most likely to reduce the risk of Lyme disease.

Most ticks start out in leaf litter or soil on the ground. When they are ready to feed, which most do three times in their life, they might crawl on top of a piece of grass and wait for an unsuspecting host.

Daniel W. Rahn, M.D.
Medical College of Georgia
Augusta, GA 30907
http://www.mcg.edu

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Grasshoppers

Phillip A. Glogoza, Extension Entomologist
Michael J. Weiss
, Professor of Entomology

Each season grasshoppers are a threat to field crops, forage crops, pastures and rangeland..

The most severe infestations are likely to occur during seasons when the weather is hot and dry.

Scouting should begin in May and early June, and producers should be prepared to start management measures when young hopper populations reach threatening levels.

Life Cycle

Grasshopper eggs are laid beneath the soil surface in pod-like structures that the female deposits from her abdomen. Each egg pod consists of 20 to 120 elongated eggs securely cemented together; the whole mass is somewhat egg shaped and covered with soil. A female grasshopper produces from eight to 25 egg masses. The species of grasshoppers that cause major crop loss overwinter in the egg stage, although a few other noneconomic species overwinter as nymphs.

In the Northern States, grasshopper egg hatch normally begins in late April to early May. The peak hatch occurs about mid June and the hatch is usually nearing completion by late June. Cool and extremely dry springs may delay the hatch, allowing it to continue into July.

Young grasshoppers are referred to as nymphs. They are similar to adults in general appearance but are smaller and have wing pads instead of wings. There are usually five or six nymphal stages and the length of time from egg to adult is 40 to 60 days. Knowledge of grasshopper instar identification is useful because it gives a rough indication of how far the hatch has progressed.

Normally, once fourth and fifth instar grasshoppers are present, the hatch is winding down. More important, recognition of fifth instar hoppers indicates that the winged adult stage is soon to follow. Winged adults are much more mobile than the nymphal stages. Wingpads of first to third instar hoppers are borne saddle-like over the thorax. Wingpads of fourth and fifth instar hoppers are pointed backward over the abdomen and differ only in size. In the fourth instar they are relatively small and extend only to the first abdominal segment, while in the fifth instar they are large and extend past the second abdominal segment.

Adults of crop-damaging grasshopper species become numerous in mid July with egg laying activity usually beginning in late July and continuing into fall. Eggs are deposited in a variety of non-crop areas including ditches, fence rows, shelterbelts and weedy areas. They are also laid in cropped areas including late season crops, weedy fallow fields and headlands as well as in haylands and alfalfa. Migratory and clear winged grasshoppers frequently lay eggs in pastureland.

Damage

Weather is the main factor affecting grasshopper population levels. Outbreaks are usually preceded by several years of hot, dry summers and warm falls, allowing populations to increase slowly.

How weather affects grasshoppers

Temperature Effects
  • High temperatures in summer - fall
        Early maturity of grasshoppers
        Long egg laying period
  • Warm spring
        Early hatch, followed by:
        <70o -->No feeding, high mortality
        Warm and dry --> Good start for
         hoppers
  • Winter temperatures have little affect
Rainfall Effects
  • Cloudy, wet weather for 1+ weeks
        Promotes fungal pathogens
        of grasshoppers
        Prolonged wet period important
  • Heavy rains during emergence
        Kills young grasshoppers
           embeds young hoppers in soil
           physically wash them away
  • Extreme drought
        Poor egg hatch
        Hoppers starve from lack of food
        Low egg production by adults

Weather effects and their impact on grasshopper populations

Decrease when . . .
  • Warm early spring
          premature hatch
          IF get a cold snap --> poor
          development
  • Hot period in early spring...
           promotes hatching
      ...following by cloudy, wet weather
           favors the occurrence of disease
  • Cool summer and early fall
           delays the maturity of the
           grasshoppers
           shortens the time for egg laying
Increase when . . .
  • Cool, wet weather in early spring
           prevents premature hatch
           insures adequate food supply
  • Warm and dry in late spring
          promotes uniform hatching time
          good weather conditions for
          feeding
  • Hot summer with adequate rainfall
          provides good food supply
          low incidence of disease
  • Late fall
          long egg laying period

Grasshopper damage to wheat and other cereal crops is generally concentrated near field margins. Individual plants will exhibit leaf stripping, beard loss after heading, head clipping, and kernels that have been fed upon or completely destroyed. When grasshopper populations are extremely high and food plants are scarce, grasshoppers migrate and will consume almost any plant they come upon. Row crop producers should be aware of the potential for grasshoppers to move into row crops after small grains have begun to dry down.

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Grubs

Knowing the life cycle of grubs is the key to determining whether you have a problem, what to do about it, and when to do it. A white grub is the immature (larval) form of a scarab beetle, such as a European chafer or Japanese beetle. Grubs live in the soil, feeding on plant roots, so you may not be aware of them until you see damage. By considering a grub’s life cycle, you can anticipate problems before your lawn is ruined. The biology of the Japanese beetle is typical of most grubs encountered in New York State and is explained below.

European chafer beetle (adult)

Adult Japanese beetle

Life cycle of Japanese beetle.

A. In late June and early July, Japanese beetle adults emerge from the ground and begin to search for food and mates. The adults can fly as far as a mile and feed on a multitude of plants; their favorites include roses, grapes, and linden trees. Other scarab beetles may go unnoticed at this time because they are not attacking ornamental plants.

B. In July, female beetles spend 2–3 weeks laying up to 60 eggs in the soil. Depending on soil moisture and temperature, eggs hatch about 2 weeks later. These first-stage ("first-instar ") grubs feed on grass roots for most of August. The grubs are small, feeding close to the surface, and vulnerable to biological and chemical insecticides at this time. Control high populations at this stage, before feeding on turf roots is noticeable.

Eggs and newly hatched grubs of Japanese beetle.

Grub larva, third stage (instar)

Pupa (becoming an adult)

 

C. From late August through October (depending on your climate), grubs molt into a second and then a third stage. As they grow, grubs consume more roots. Damaged turf often appears now.

D. As temperatures drop in autumn, grubs move down in the soil. They overwinter as third-instar grubs below the frost line.

E. In the spring, they move up in the soil to feed on roots for a very short time. (Most of the lawn damage seen in the spring is a result of fall feeding, not spring feeding.)

F. In late spring, grubs stop feeding and turn into pupae that are resistant to insecticides. In late June or early July, beetles emerge from the pupae and crawl out of the soil, completing the cycle.

THE GRUB-DAMAGED LAWN

Severe grub damage in a lawn appears as large, irregular sections of brown turf that detach from the soil without effort. Unlike turf damaged by drought or excessive fertilizer, the turf peels away like a carpet being rolled up.

For most of the year, however, grubs are out of sight and out of mind. They feed on grass roots in your lawn and are usually noticed only when dead and damaged areas appear.

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Green June Beetle

The green June beetle is in the Scarabaieidae family and also referred to as white grubs. Unlike many of the other white grub pest of turfgrass this species is unique in that it will come to the soil surface and crawl on the turfgrass at night. Their larval tunneling activity can damage turfgrass stands.

Identification

The adult green June beetle is usually 3/4" to 1" long, and 1/2" wide. The top side is forest green, with or without lengthwise tan stripes on the wings. The underside is metallic bright green or gold, bearing legs with stout spines to aid in digging. In the Mid-Atlantic region the names "June bug" and "June beetle" are commonly used for this insect. They're called "fig eater" in the southern part of their range. Do not confuse the green June beetle, however, with the familiar brown May or June beetles that are seen flying to lights on summer nights. The green June beetle adult flies only during the day.

The larvae are white grubs often called "Richworms" because they prefer "high" levels of organic matter for food. With three growth stages they develop and are similar to the other annual scarab species. Their body lengths reach 1/4", 3/4", and 2" respectively. The larvae have stiff abdominal bristles, short stubby legs, and wide body. One unique characteristic of this grub is that it crawls on its back by undulating and utilizing its abdominal bristles to gain traction. Other typical white grubs, like the Japanese beetle grub, are narrower, have longer legs, crawl right side up and when at rest assume a "C" shaped posture.

The most closely related U. S. species is Cotinis mutablis.

Distribution

This species is native to the eastern half of the United States and overlaps with Cotinis mutablis in Texas and the southwestern United States.

Hosts

The adults generally don't feed but occasionally become a pest of fruit. Any thin skin fruit such as fig, peach, plum, blackberry, grape and apricot can be eaten. The principal attraction is probably the moisture and the fermenting sugars of ripening fruit. They occasionally feed on plant sap. In turf situations egg laying females are attracted to moist sandy soils with high levels of organic matter. Turf areas treated repeatedly with organic fertilizers, composts or composted sewage sludge become more attractive to the female.

The grub feeds on dead, decaying organic matter as well as plant roots. This species is commonly associated with both agricultural crop and livestock production areas as well as urban landscapes. Field stored hay bales, manure piles, grass clipping piles, bark mulches and other sources of plant material that come in contact with moist soil are prime microhabitats preferred by both the female for egg laying and the migrating 3rd instar grubs

Life Cycle

The green June beetle completes one generation each year. Adults begin flying in June and may continue sporadically into September. The peak occurrence of adults is during a two week period in mid-July in Maryland and Virginia. On warm sunny days, adults may swarm over open grassy areas. Their flight behavior and sounds resembles that of a bumble bee. At night they rest in trees or beneath the thatch.

The adult females shortly after emerging may fly to the lower limbs of trees and shrubs and release a pheromone that attracts large numbers of males. Frequently, males repeatedly fly low and erratic over the turf trying to locate emerging females. After mating, females burrow 2" to 8" into the soil to lay about twenty eggs at a time. The spherical eggs are white and almost 1/16" in diameter.

Most eggs hatch in late July and August. The first two instar stages feed at the soil thatch interface. By the end of September, most are third instar larvae and these large grubs tunnel into the thatch layer and construct a deep vertical burrow. The grubs may remain active into November in the Mid-Atlantic region. In the more southern states grubs may become active on warm nights throughout the winter. In colder areas they overwinter in burrows 8"-30" deep. The grubs resume feeding once the ground warms in the spring and pupate in late May or early June. The adults begin emerging about three weeks later.

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Leafhoppers

Leafhoppers are one of the largest families of plant-feeding insects. There are more leafhopper species worldwide than all species of birds, mammals, reptiles, and amphibians combined. Leafhoppers feed by sucking the sap of vascular plants, and are found almost anywhere such plants occur, from tropical rainforests, to arctic tundra. Several leafhopper species are important agricultural pests.

Details of the life cycle vary from species to species. In general, the female inserts several eggs into the living tissue of the host plant. The eggs either remain dormant for a period ranging from a month to over a year, or develop and hatch within a few weeks. The young, known as nymphs, feed on plant sap by inserting their beaks into the vascular or parenchyma tissues of the host plant and go through a series of five moults (shedding their exoskeleton), reaching the adult stage after a period of several weeks or months. Adult males and females seek each other out for mating, locating each other through specialized courtship calls.

All feed on plant sap. Leafhopper species feed on a wide variety of vascular plant species, including grasses, sedges, broad-leafed woody and herbaceous plants of many families, and conifers. At least one leafhopper species can usually be found feeding on the each of the dominant plant species in practically every terrestrial ecosystem. Frequently several leafhopper species can be found coexisting on the same plant.

Nationwide, the potato leafhopper is a very injurious pest of forages, particularly alfalfa and clover. Both nymphs and adults feed on the undersides of the leaves. By extracting the sap, they cause stunting and leaf curl, as well as the condition called "hopperburn." This disease is caused by the injection of a toxic substance. It is characterized by a yellowing of the tissue at the tip and around the leaf margin which increases until the leaf dies. Symptoms are sometimes confused with drought stress.

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Spittlebug 

The Saratoga spittlebug, Aphrophora saratogensis (Fitch)2, so called because it was first collected in SaratogaCover Photo County, N. Y., is a native insect that is destructive to several species of pine in Eastern North America. It occurs where ever its host grows, from Maine to Minnesota in the United States and in the southern portions of the adjacent Canadian Provinces.

The adult of this insect often destroys young pines, especially in plantations where its alternate hosts are abundant. Natural-grown and large trees usually are less injured.

Hosts
Figure 1
Figure 1 - Sweet-fern plant-the principal host of the spittlebug nymph.
Red pine is the preferred host of the adult spittlebug. Jack pine follows, although decreased planting of this species in recent years has lessened its importance as a host. Scots pine, which is increasingly planted for Christmas trees, is occasionally injured by the spittlebug.

White pine is frequently fed upon but seldom damaged severely. Adult spittlebugs thought to be Aphrophora saratogensis have been collected from pitch pine, tamarack, balsam fir, and northern white-cedar-usually from trees near infested red pine. The nymphs require two alternate hosts for their development. The early stages or instars feed on herbaceous species of plants of the forest floor such as brambles (raspberry and blackberry), orange hawkweed, everlasting, aster, and many others. Older nymphs feed on sweet-fern (fig. 1) and willow sprouts.

Damage